CODEXIS, INC. - Common Stock, par value $0.0001 per share (CDXS) Institutional Ownership

13F Institutional Holders and Ownership History from Q1 2014 to Q4 2025

Type / Class
Equity / Common Stock, par value $0.0001 per share
Symbol
CDXS on Nasdaq
Shares outstanding
90,289,069
Price per share
$1.30
All holders as of 31 Dec 2025
Q4 2025
Total 13F shares
64,649,771
Total reported value
$105,376,439
% of total 13F portfolios
0%
Share change
-1,490,751
Value change
-$6,330,751
Number of holders
123
Price from insider filings
$1.30
Reported Shares, Excluding Options vs Price
Reported Value, Excluding Options
Total Investors

Significant Owners of CODEXIS, INC. - Common Stock, par value $0.0001 per share (CDXS) based on Schedule 13D/G filings

Filer Ownership Change % Holdings Value Net Change Nb Shares Change % Reporting Name Report Period
Aberdeen Group plc 8.8% +86% $12,985,392 +$6,315,331 7,966,498 +95% Aberdeen Group plc 31 Dec 2025
Opaleye Management Inc. 7.5% -24% $11,067,273 -$3,583,149 6,789,738 -24% Opaleye Management Inc. 31 Dec 2025
BlackRock, Inc. 7.2% $13,752,804 5,979,480 BlackRock, Inc. 31 Mar 2025
Nantahala Capital Management, LLC 4.3% $6,825,958 2,967,808 Nantahala Capital Management, LLC 30 Sep 2024
FMR LLC 0.6% $1,425,855 584,367 FMR LLC 30 Sep 2025

As of 31 Dec 2025, 123 institutional investors reported holding 64,649,771 shares of CODEXIS, INC. - Common Stock, par value $0.0001 per share (CDXS). This represents 72% of the company’s total 90,289,069 outstanding shares.

Top 25 Institutional Shareholders

The largest institutional shareholders of CODEXIS, INC. - Common Stock, par value $0.0001 per share (CDXS) together control 64% of the company’s outstanding shares — a level of ownership often associated with strong institutional conviction and long‑term capital commitment.

Institution Ownership % Shares Held Share Change % Portfolio % Holdings Value
Aberdeen Group plc 8.8% 7,966,498 +95% 0.02% $12,985,392
BlackRock, Inc. 8.2% 7,395,740 +3.5% 0% $12,055,056
Opaleye Management Inc. 7.5% 6,789,738 -47% 1.5% $11,067,273
VANGUARD GROUP INC 5.5% 4,941,542 +5.8% 0% $8,054,714
Nantahala Capital Management, LLC 4.4% 3,990,169 -1% 0.39% $6,503,975
Telemark Asset Management, LLC 4.3% 3,846,411 -3.8% 0.63% $6,269,650
AMERIPRISE FINANCIAL INC 4.2% 3,793,574 +37% 0% $6,183,525
MILLENNIUM MANAGEMENT LLC 3.5% 3,135,192 -23% 0% $5,110,363
Prescott Group Capital Management, L.L.C. 2.8% 2,568,359 +304% 0.39% $4,186,425
GEODE CAPITAL MANAGEMENT, LLC 2.2% 2,026,006 +1.7% 0% $3,303,021
STATE STREET CORP 1.8% 1,603,510 +4.1% 0% $2,613,721
JACOBS LEVY EQUITY MANAGEMENT, INC 1.3% 1,135,190 -2.5% 0.01% $1,850,360
UBS Group AG 0.9% 809,520 +55% 0% $1,319,518
FRANKLIN RESOURCES INC 0.82% 740,558 -2.1% 0% $1,207,109
RENAISSANCE TECHNOLOGIES LLC 0.82% 737,485 +138% 0% $1,202,101
GOLDMAN SACHS GROUP INC 0.8% 718,109 -0.85% 0% $1,170,518
Private Advisor Group, LLC 0.72% 654,581 +17% 0.01% $1,066,967
NORTHERN TRUST CORP 0.71% 645,296 +0% 0% $1,051,833
FMR LLC 0.71% 641,926 +9.8% 0% $1,046,340
D. E. Shaw & Co., Inc. 0.7% 628,291 +261% 0% $1,024,114
Assenagon Asset Management S.A. 0.63% 566,547 +142% 0% $923,472
TUDOR INVESTMENT CORP ET AL 0.6% 537,833 0% 0% $876,668
PERKINS CAPITAL MANAGEMENT INC 0.59% 529,500 +72% 0.69% $863,085
Post Resch Tallon Group Inc. 0.57% 511,748 -8.6% 0.7% $834,149
CITADEL ADVISORS LLC 0.55% 499,377 +85% 0% $813,985

Institutional Holders of CODEXIS, INC. - Common Stock, par value $0.0001 per share (CDXS) across Reporting Periods

Period Reported Shares, Excl. Options Value, Excl. Options Value Change Price (Median) Investors
2025 Q4 64,649,771 $105,376,439 -$6,330,751 $1.63 123
2025 Q3 65,969,954 $160,958,513 -$12,809,765 $2.44 129
2025 Q2 71,218,652 $173,772,833 +$14,555,678 $2.44 117
2025 Q1 65,178,959 $175,225,389 +$929,989 $2.69 124
2024 Q4 63,988,678 $305,316,642 +$3,089,141 $4.77 133
2024 Q3 62,919,076 $193,796,579 +$32,967,849 $3.08 115
2024 Q2 52,477,224 $162,676,431 -$4,430,246 $3.10 117
2024 Q1 53,718,149 $187,413,304 -$5,510,895 $3.49 123
2023 Q4 55,230,099 $168,449,151 +$3,188,960 $3.05 117
2023 Q3 55,050,956 $104,041,277 -$17,760,919 $1.89 119
2023 Q2 60,904,680 $170,515,502 +$1,545,759 $2.80 122
2023 Q1 59,089,040 $244,372,878 -$3,021,145 $4.14 144
2022 Q4 59,636,802 $788,383,480 -$13,405,240 $4.66 161
2022 Q3 61,301,294 $371,491,238 -$40,070,138 $6.06 164
2022 Q2 64,991,812 $680,516,879 -$6,800,772 $10.46 162
2022 Q1 61,863,148 $1,274,964,586 -$98,934,303 $20.62 151
2021 Q4 67,147,021 $2,095,342,832 +$76,348,853 $31.27 170
2021 Q3 63,138,366 $1,471,997,186 -$5,907,120 $23.26 152
2021 Q2 63,445,130 $1,435,375,799 +$28,573,787 $22.66 141
2021 Q1 62,157,382 $1,422,750,683 +$27,881,773 $22.89 150
2020 Q4 61,031,096 $1,332,428,797 +$147,547,692 $21.83 149
2020 Q3 54,143,314 $635,384,057 +$756,055 $11.74 125
2020 Q2 54,090,083 $615,832,306 -$9,739,461 $11.40 123
2020 Q1 54,973,182 $613,470,073 +$35,167,629 $11.16 116
2019 Q4 51,136,893 $817,632,585 +$18,855,885 $15.99 120
2019 Q3 49,670,282 $681,242,929 -$37,267,524 $13.72 116
2019 Q2 50,745,755 $928,253,177 +$71,892,883 $18.43 121
2019 Q1 46,639,678 $957,504,084 +$11,137,897 $20.53 122
2018 Q4 46,184,102 $771,318,098 +$46,246,024 $16.70 119
2018 Q3 43,408,808 $744,445,459 +$36,964,971 $17.15 108
2018 Q2 41,280,432 $594,459,182 +$90,212,376 $14.40 99
2018 Q1 35,090,414 $385,990,722 +$11,240,688 $11.00 79
2017 Q4 34,306,430 $286,457,460 +$20,633,224 $8.35 73
2017 Q3 31,764,185 $211,226,933 +$6,875,107 $6.65 62
2017 Q2 30,740,171 $167,534,547 +$33,321,129 $5.45 56
2017 Q1 24,570,510 $117,934,000 +$14,219,009 $4.80 55
2016 Q4 24,154,209 $111,107,286 +$2,230,034 $4.60 58
2016 Q3 23,606,886 $104,808,783 +$180,655 $4.44 60
2016 Q2 23,568,734 $94,983,002 +$14,252,595 $4.03 55
2016 Q1 20,217,995 $62,878,000 +$2,277,261 $3.11 50
2015 Q4 13,854,689 $58,596,699 +$3,498,720 $4.23 46
2015 Q3 13,064,117 $44,279,501 +$2,421,990 $3.39 46
2015 Q2 12,291,372 $47,805,344 +$6,217,663 $3.89 50
2015 Q1 10,707,704 $48,822,000 +$5,406,301 $4.56 40
2014 Q4 9,767,724 $24,611,000 -$2,044,811 $2.52 33
2014 Q3 10,622,351 $24,528,700 +$1,600,542 $2.33 33
2014 Q2 10,560,796 $15,419,009 +$324,151 $1.46 31
2014 Q1 10,329,923 $21,081,175 +$2,030,404 $2.04 34