Comparable stake leader
Aberdeen Group plc
SEC ownership of CODEXIS, INC.
Top holders, 13D/G large-stake signals, insider positions, and transaction flow tied to CIK 0001200375.
Disclosed value leader
Opaleye Management Inc.
Comparable rows
50/50
Latest evidence
11 Jun 2026
- Public ticker
- CDXS on Nasdaq
- Parsed ownership sources
- 13F, 13D/G, and 3/4/5
- Largest SEC-reported holder
- Aberdeen Group plc
- Latest 13D/G holder
- Aberdeen Group plc
- Latest parsed ownership date
- 11 Jun 2026
- Issuer CIK
- 0001200375
- Issuer location
- Redwood City, CA
Ownership snapshot
CIK-linked ownership evidence from parsed SEC filings.
2 signals
Issuer
Names, identifiers, and filing context for this profile.
2 identity fields
Ownership snapshot
Top shareholders of CODEXIS, INC. (13F + 13D/G + 3/4/5)
Grouped by holder across the latest SEC ownership disclosures. Start with comparable ownership when available, then verify the linked 13F, 13D/G, or 3/4/5 evidence.
Comparable stake leader
Aberdeen Group plc
Disclosed value leader
Opaleye Management Inc.
Comparable rows
50/50
Latest evidence
11 Jun 2026
13F
13D/G
3/4/5
Lead comparable stake: 8.8%
13D/G row: Aberdeen Group plc
Showing 1-8 of 50 holder rows.
Quick read
Aberdeen Group plc leads the comparable SEC ownership view at 8.8%.
50 of 50 rows have a safely comparable ownership percentage. Use this as a filing-backed starting point, then open the detailed table for source links, reporting role, position value, share count, and filing date.
Open SEC evidence
Opens Aberdeen Group plc's linked filing trail.
Comparable ownership
Top 5
Aberdeen Group plc
8.8%
Opaleye Management Inc.
7.5%
BlackRock, Inc.
7.2%
INTEGRATED CORE STRATEGIES (US) LLC
5.2%
Laurence W. Lytton
5%
Ownership % is shown only when SEC disclosures are safely comparable across share classes.
Detailed SEC evidence table Showing 1-8 of 50 holder rows
| Holder | Evidence | Ownership % | Position | 12M Net Δ | As of |
|---|---|---|---|---|---|
| Aberdeen Group plc |
13F
13D/G
Company
|
8.8%
from 13D/G
|
$14,149,000
8,680,368 shares
|
— | 31 Mar 2026 |
| Opaleye Management Inc. |
3/4/5
13D/G
13F
10%+ Owner · Company
|
7.5%
from 13D/G
|
$16,900,093
10,134,989 shares
|
-$507,767 | 26 Nov 2025 |
| BlackRock, Inc. |
13D/G
13F
Company
|
7.2%
|
$13,752,804
5,979,480 shares
|
$0 | 31 Mar 2025 |
| INTEGRATED CORE STRATEGIES (US) LLC |
13D/G
|
5.2%
|
$7,812,135
4,734,627 shares
|
$0 | 09 Apr 2026 |
| Laurence W. Lytton |
13D/G
LYTTON LAURENCE W
|
5%
|
$5,347,841
4,532,069 shares
|
$0 | 06 Mar 2026 |
| Nantahala Capital Management, LLC |
13D/G
13F
Company
|
4.3%
|
$6,825,958
2,967,808 shares
|
$0 | 30 Sep 2024 |
| FMR LLC |
13D/G
13F
Company
|
0.6%
|
$1,425,855
584,367 shares
|
$0 | 30 Sep 2025 |
| David V Smith |
3/4/5
Director
|
0.21%
|
$489,572
194,275 shares
|
— | 10 Jun 2025 |
How to read this table
Rows are grouped by holder and ranked first by SEC-reported ownership % when it is safely comparable, then by disclosed holdings value. — indicates unavailable or non-comparable values; "mixed-class rows" means the latest filing spans non-comparable classes.
Forms 3/4/5 flow
Insider trading overview
Net insider flow leans to selling over the last 12 months. Net value: -$368,498.
Buys
$2,836,198
Insiders: 1
Sells
$3,204,697
Insiders: 6
Net
-$368,498
Shares: -481,150
Window
Last 12 months
4 rolling buckets
Buy value
Sell value
Shares: 1,335,000
Shares: 1,816,150
| Window (months) | Buy shares | Sell shares | Buy value | Sell value | Net value |
|---|---|---|---|---|---|
| 0-3 | 0 | 2,605 | $0 | $6,174 | -$6,174 |
| 3-6 | 0 | 20,691 | $0 | $27,850 | -$27,850 |
| 6-9 | 890,000 | 1,792,854 | $1,724,108 | $3,170,673 | -$1,446,565 |
| 9-12 | 445,000 | 0 | $1,112,090 | $0 | +$1,112,090 |
Reporting owners
Insider holders from Forms 3/4/5
Showing 1-20 of 22 reporting-owner rows connected to CODEXIS, INC.. Open a relationship link for the CIK-to-CIK filing trail.
Rows
22
Largest Forms 3/4/5 holder
Opaleye Management Inc.
Net 12M change
-$589,495
Latest report
11 Jun 2026
| Name | Relationship | Holdings Value | Past Year Net Change | Change % | Valuation | Report Date |
|---|---|---|---|---|---|---|
| Opaleye Management Inc. | 10%+ Owner | $16,900,093 | -$507,767 | -2.9% | Filing P/S | 26 Nov 2025 |
| John J. Nicols | Director | $1,652,912 | Mixed | 17 Apr 2023 | ||
| Stephen G. Dilly | Director | $1,343,085 | Mixed | 11 Nov 2025 | ||
| Kevin Norrett | Chief Operating Officer | $761,265 | Mixed | 05 Feb 2025 | ||
| David V Smith | Director | $489,572 | Mixed | 10 Jun 2025 | ||
| Byron L. Dorgan | Director | $471,398 | Mixed | 11 Jun 2024 | ||
| Bernard J. Kelley | Director | $434,617 | Mixed | 10 Aug 2021 | ||
| Sriram Ryali | Chief Financial Officer | $370,638 | Mixed | 01 Jan 2024 | ||
| PARKER H STEWART | Director | $361,829 | Mixed | 10 Jun 2025 | ||
| Rahul Singhvi | Director | $345,371 | Mixed | 10 Jun 2025 | ||
| Alison Moore | President and CEO, Director | $327,436 | -$20,630 | -5.9% | Filing P/S | 23 Feb 2026 |
| Esther Martinborough | Director | $325,170 | Mixed | 10 Jun 2025 | ||
| Cynthia Collins | Director | $282,635 | Mixed | 10 Jun 2025 | ||
| Margaret Nell Fitzgerald | Chief Legal and Compliance Officer | $282,041 | Mixed | 01 Jan 2024 | ||
| Dennis P. Wolf | Director | $217,124 | -$13,988 | -6.1% | Filing P/S | 21 Nov 2025 |
| Christos Richards | Director | $214,535 | Mixed | 10 Jun 2025 | ||
| Raymond de Vre | Director | $201,002 | -$8,632 | -4.1% | Filing P/S | 11 Jun 2026 |
| Stefan Lutz | Chief Scientific Officer | $147,753 | -$17,848 | -11% | Filing P/S | 23 Feb 2026 |
| Jennifer Aaker | Director | $139,599 | Mixed | 13 Jun 2023 | ||
| Patrick Y. Yang | Director | $113,400 | Mixed | 14 Jun 2022 |
Recent insider transactions for CODEXIS, INC.
Showing up to 300 recent insider transactions from the last 12 months.
| Insider | Symbol | Class | Transaction | % | SEC-reported value | SEC-reported price | Shares | Shares owned | Date | Ownership |
|---|---|---|---|---|---|---|---|---|---|---|
| Raymond de Vre | CDXS | Common Stock | Sale | -2.98% | $6,174 | $2.37 | -2,605 | 84,811 | 11 Jun 2026 | Direct |
| Stefan Lutz | CDXS | Common Stock | Sale | -4.3% | $6,641 | $1.13 | -5,862 | 130,421 | 24 Feb 2026 | Direct |
| Stefan Lutz | CDXS | Stock Option (Right to Buy) | Award | 316,100 | 316,100 | 23 Feb 2026 | Direct | |||
| Alison Moore | CDXS | Stock Option (Right to Buy) | Award | 993,000 | 993,000 | 23 Feb 2026 | Direct | |||
| Georgia Erbez | CDXS | Stock Option (Right to Buy) | Award | 316,100 | 316,100 | 23 Feb 2026 | Direct | |||
| Stefan Lutz | CDXS | Common Stock | Sale | -1.8% | $3,257 | $1.31 | -2,492 | 136,283 | 06 Feb 2026 | Direct |
| Georgia Erbez | CDXS | Common Stock | Sale | -5.42% | $5,001 | $1.31 | -3,826 | 66,810 | 06 Feb 2026 | Direct |
| Alison Moore | CDXS | Common Stock | Sale | -1.5% | $5,001 | $1.31 | -3,826 | 250,506 | 06 Feb 2026 | Direct |
| Stefan Lutz | CDXS | Common Stock | Sale | -3.27% | $7,950 | $1.70 | -4,685 | 138,775 | 07 Jan 2026 | Direct |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -4.53% | $14,229 | $1.67 | -8,533 | 179,989 | 01 Dec 2025 | By Managed Account |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -2.4% | $408,538 | $1.67 | -245,000 | 9,955,000 | 01 Dec 2025 | By Opaleye, L.P. |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -1.3% | $4,358 | $1.75 | -2,490 | 188,522 | 28 Nov 2025 | By Managed Account |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -1.02% | $183,750 | $1.75 | -105,000 | 10,200,000 | 28 Nov 2025 | By Opaleye, L.P. |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -2.55% | $8,706 | $1.74 | -5,000 | 191,012 | 26 Nov 2025 | By Managed Account |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -1.42% | $258,266 | $1.74 | -148,318 | 10,305,000 | 26 Nov 2025 | By Opaleye, L.P. |
| Dennis P. Wolf | CDXS | Common Stock | Sale | -6.05% | $13,988 | $1.56 | -8,952 | 138,951 | 21 Nov 2025 | Direct |
| Alison Moore | CDXS | Common Stock | Award | 50.2% | 85,000 | 254,332 | 13 Nov 2025 | Direct | ||
| Alison Moore | CDXS | Stock Option (Right to Buy) | Award | 500,000 | 500,000 | 13 Nov 2025 | Direct | |||
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -1.55% | $323,730 | $1.96 | -165,000 | 10,453,318 | 12 Nov 2025 | By Opaleye, L.P. |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -12.3% | $53,194 | $1.94 | -27,459 | 196,012 | 12 Nov 2025 | By Managed Account |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | 9.15% | $1,724,108 | $1.94 | 890,000 | 10,618,318 | 12 Nov 2025 | By Opaleye, L.P. |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -5.58% | $23,772 | $1.80 | -13,211 | 223,461 | 11 Nov 2025 | By Managed Account |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -4.44% | $962,679 | $1.80 | -535,000 | 11,508,318 | 11 Nov 2025 | By Opaleye, L.P. |
| Raymond de Vre | CDXS | Common Stock | Sale | -1.59% | $2,459 | $1.74 | -1,413 | 87,416 | 11 Nov 2025 | Direct |
| Stephen G. Dilly | CDXS | Common Stock | Award | 7.38% | 50,000 | 727,053 | 11 Nov 2025 | Direct | ||
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -1.38% | $5,756 | $1.73 | -3,318 | 236,682 | 10 Nov 2025 | By Managed Account |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -1.67% | $355,614 | $1.73 | -205,000 | 12,043,318 | 10 Nov 2025 | By Opaleye, L.P. |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -4% | $16,968 | $1.70 | -10,000 | 240,000 | 07 Nov 2025 | By Managed Account |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Sale | -2.36% | $503,410 | $1.70 | -296,682 | 12,248,318 | 07 Nov 2025 | By Opaleye, L.P. |
| Alison Moore | CDXS | Common Stock | Sale | -3.55% | $15,629 | $2.51 | -6,239 | 169,332 | 13 Oct 2025 | Direct |
| Georgia Erbez | CDXS | Common Stock | Sale | -8.12% | $15,629 | $2.50 | -6,239 | 70,636 | 13 Oct 2025 | Direct |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Purchase | 0.01% | $2,211 | $2.84 | 780 | 12,545,000 | 04 Aug 2025 | By Opaleye, L.P. |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Purchase | 0.68% | $225,710 | $2.68 | 84,220 | 12,544,220 | 31 Jul 2025 | By Opaleye, L.P. |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Purchase | 0.01% | $4,411 | $2.80 | 1,578 | 12,460,000 | 30 Jul 2025 | By Opaleye, L.P. |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Purchase | 0.07% | $23,750 | $2.82 | 8,422 | 12,458,422 | 29 Jul 2025 | By Opaleye, L.P. |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Purchase | 0.81% | $282,000 | $2.82 | 100,000 | 12,450,000 | 18 Jul 2025 | By Opaleye, L.P. |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Purchase | 0.82% | $239,000 | $2.39 | 100,000 | 12,350,000 | 27 Jun 2025 | By Opaleye, L.P. |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Purchase | 0.42% | $114,012 | $2.24 | 50,898 | 12,250,000 | 16 Jun 2025 | By Opaleye, L.P. |
| Opaleye Management Inc. | CDXS | Common Stock, par value $0.001 per share | Purchase | 0.82% | $220,997 | $2.23 | 99,102 | 12,199,102 | 13 Jun 2025 | By Opaleye, L.P. |
* marks a reported price that did not pass the local price check.