| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| JANUS HENDERSON GROUP PLC | 12.4% | 2.53M | 0 | 2.53M | Kristin Mariani | Head of North America Compliance, CCO | |
| Janus Henderson Biotech Innovation Master Fund Ltd | 5.8% | 1.19M | 0 | 1.19M | Kristin Mariani | Head of North America Compliance, CCO |