| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| Perceptive Advisors LLC | 13.7% | 540,296 | 0 | 540,296 | /s/ Joseph Edelman | Joseph Edelman, Managing Member | 0001224962 |
| Joseph Edelman | 13.9% | 550,024 | 0 | 550,024 | /s/ Joseph Edelman | Joseph Edelman | 0001164426 |
| Perceptive Life Sciences Master Fund, Ltd. | 13.7% | 540,296 | 0 | 540,296 | /s/ Joseph Edelman | Joseph Edelman, Managing Member | 0001249675 |