| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| JANUS HENDERSON GROUP PLC | 16.3% | 1,145,660 | 0 | 1,145,660 | Kristin Mariani | Head of North America Compliance, CCO | |
| Janus Henderson Biotech Innovation Master Fund Ltd | 14% | 984,485 | 0 | 984,485 | Kristin Mariani | Head of North America Compliance, CCO |