| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| JANUS HENDERSON GROUP PLC | 18.6% | 1.13M | 0 | 1.13M | Kristin Mariani | Head of North America Compliance, CCO | |
| Janus Henderson Biotech Innovation Master Fund Ltd | 16.5% | 1.01M | 0 | 1.01M | Kristin Mariani | Head of North America Compliance, CCO |