| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| JANUS HENDERSON GROUP PLC | 17.2% | 1,046,899 | 0 | 1,046,899 | Kristin Mariani | Head of North America Compliance, CCO | |
| Janus Henderson Biotech Innovation Master Fund Ltd | 15.6% | 952,829 | 0 | 952,829 | Kristin Mariani | Head of North America Compliance, CCO |