| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| JANUS HENDERSON GROUP PLC | 11.6% | 804,976 | 0 | 804,976 | Kristin Mariani | Head of North America Compliance, CCO | |
| Janus Henderson Biotech Innovation Master Fund Ltd | 10.4% | 719,574 | 0 | 719,574 | Kristin Mariani | Head of North America Compliance, CCO |