| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| OrbiMed Advisors LLC | 8.9% | 967K | 0 | 967K | /s/ Carl L. Gordon | Carl L. Gordon / Member of OrbiMed Advisors LLC | |
| OrbiMed Capital GP IX LLC | 6% | 645K | 0 | 645K | /s/ Carl L. Gordon | Carl L. Gordon / Member |