| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| OrbiMed Advisors LLC | 7.2% | 2,133,405 | 0 | 2,133,405 | /s/ Carl L. Gordon | Carl L. Gordon / Member | |
| OrbiMed Capital GP IX LLC | 6.3% | 1,878,704 | 0 | 1,878,704 | /s/ Carl L. Gordon | Carl L. Gordon / Member of OrbiMed Advisors LLC |