Aberdeen Group plc - 01 Oct 2025 SCHEDULE 13G/A Report for CODEXIS, INC. Common Stock, par value $0.0001 per share (CDXS)

Filing Manager: Aberdeen Group plc
Reporting Manager: Aberdeen Group plc
Security: CODEXIS, INC. Common Stock, par value $0.0001 per share
Symbol: CDXS
Shares outstanding: 90.3M shares
Disclosed Ownership: 4.09M shares
Ownership: 4.53%
Form type: SCHEDULE 13G/A
Filing time: 07 Nov 2025, 11:28:10 UTC
Date of event: 01 Oct 2025
Previous filing: 30 Jul 2025
SEC: View on sec.gov

Reporting Managers (2)

Name	Ownership	Shares Owned	sole voting power	shared voting power	Signature	Title	CIK
Aberdeen Group plc	4.53%	4.09M	0	4.09M	Jenny Clarke	Major Shareholding Reporting Analyst
abrdn Inc.	4.53%	4.09M	0	4.09M	Jenny Clarke	Major Shareholding Reporting Analyst