| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| Lundbeckfond Invest A/S | 1.6% | 203,122 | 203,122 | 0 | /s/ Lene Skole | By Lene Skole, Chief Executive Officer & Attorney-in-Fact | |
| Lene Skole | 1.6% | 203,122 | 203,122 | 0 | /s/ Lene Skole | Lene Skole |