| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| JANUS HENDERSON GROUP PLC | 20% | 1,287,368 | 0 | 1,287,368 | Kristin Mariani | Head of North America Compliance, CCO | |
| Janus Henderson Biotech Innovation Master Fund Ltd | 17% | 1,105,471 | 0 | 1,105,471 | Kristin Mariani | Head of North America Compliance, CCO |