| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| Ikarian Capital, LLC | 4.8% | 3,543,359 | 0 | 3,543,359 | /s/ Neil Shahrestani | Neil Shahrestani, Sole Manager | |
| Neil Shahrestani | 4.8% | 3,543,359 | 0 | 3,543,359 | /s/ Neil Shahrestani | Neil Shahrestani |