| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| Alexandria Real Estate Equities, Inc. | 4.3% | 6,500,000 | 0 | 6,500,000 | /s/ Marc E. Binda | By Marc E. Binda, Chief Financial Officer and Treasurer | |
| Alexandria Venture Investments, LLC | 3% | 4,450,000 | 0 | 4,450,000 | /s/ Marc E. Binda | By Alexandria Real Estate Equities, Inc., managing member, By Marc E. Binda, Chief Financial Officer and Treasurer | |
| Alexandria Equities No. 7, LLC | 1.4% | 2,050,000 | 0 | 2,050,000 | /s/ Marc E. Binda | See Note 1 |