| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| Biogen Inc. | 10% | 6,241,473 | 0 | 6,241,473 | /s/ Michael Dambach | Michael Dambach / Treasurer | 0000875045 |
| Biogen MA Inc. | 10% | 6,241,473 | 0 | 6,241,473 | /s/ Michael Dambach | Michael Dambach / Treasurer |