| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| OrbiMed Capital LLC | 5.1% | 2,184,200 | 2,184,200 | 0 | /s/ Carl L. Gordon | Carl L. Gordon / Member | |
| OrbiMed Advisors LLC | 0.9% | 394,779 | 0 | 394,779 | /s/ Carl L. Gordon | Carl L. Gordon / Member |