| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| OrbiMed Capital LLC | 7% | 2,969,500 | 2,969,500 | 0 | /s/ Carl L. Gordon | Carl L. Gordon / Member | |
| OrbiMed Advisors LLC | 1.1% | 473,353 | 0 | 473,353 | /s/ Carl L. Gordon | Carl L. Gordon / Member |