| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| EcoR1 Capital, LLC | 30.7% | 22,970,388 | 0 | 22,970,388 | /s/ Oleg Nodelman | Manager | 0001587114 |
| Oleg Nodelman | 30.7% | 22,970,388 | 0 | 22,970,388 | /s/ Oleg Nodelman | Reporting person | 0001454385 |
| EcoR1 Capital Fund Qualified, L.P. | 28.8% | 21,582,563 | 0 | 21,582,563 | /s/ Oleg Nodelman | Manager of the General Partner, EcoR1 Capital, LLC | 0001598292 |