| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| EcoR1 Capital, LLC | 1.3% | 662,833 | 0 | 662,833 | /s/ Oleg Nodelman | Manager | |
| Oleg Nodelman | 1.3% | 662,833 | 0 | 662,833 | /s/ Oleg Nodelman | Reporting person | |
| EcoR1 Capital Fund Qualified, L.P. | 1.2% | 623,967 | 0 | 623,967 | /s/ Oleg Nodelman | Manager of the General Partner, EcoR1 Capital, LLC |