| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| EcoR1 Capital, LLC | 1.4% | 345,253 | 0 | 345,253 | /s/ Oleg Nodelman | Manager | |
| Oleg Nodelman | 1.4% | 345,253 | 0 | 345,253 | /s/ Oleg Nodelman | Reporting person | |
| EcoR1 Capital Fund Qualified, L.P. | 1.3% | 324,365 | 0 | 324,365 | /s/ Oleg Nodelman | Manager of the General Partner, EcoR1 Capital, LLC |