| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| EcoR1 Capital, LLC | 13% | 8,714,522 | 0 | 8,714,522 | /s/ Oleg Nodelman | Manager | 0001587114 |
| Oleg Nodelman | 13% | 8,714,522 | 0 | 8,714,522 | /s/ Oleg Nodelman | Reporting Person | 0001454385 |
| EcoR1 Capital Fund Qualified, L.P. | 12% | 8,192,077 | 0 | 8,192,077 | /s/ Oleg Nodelman | Manager of the General Partner, EcoR1 Capital, LLC | 0001598292 |
| EcoR1 Capital Fund, L.P. | 0.79% | 522,445 | 0 | 522,445 | /s/ Oleg Nodelman | Manager of the General Partner, EcoR1 Capital, LLC | 0001567938 |