| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| The Column Group II, LP | 3.6% | 3.54M | 0 | 3.54M | /s/ James Evangelista, Attorney-in-Fact | James Evangelista, Attorney-in-Fact | |
| The Column Group II GP, LP | 3.6% | 3.54M | 0 | 3.54M | /s/ James Evangelista, Attorney-in-Fact | James Evangelista, Attorney-in-Fact | |
| David V Goeddel | 3.7% | 3.63M | 86.7K | 3.54M | /s/ James Evangelista, Attorney-in-Fact | James Evangelista, Attorney-in-Fact | |
| Peter Svennilson | 3.7% | 3.64M | 103K | 3.54M | /s/ James Evangelista, Attorney-in-Fact | James Evangelista, Attorney-in-Fact |