| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| Gilead Sciences, Inc. | 29.9% | 20,981,010 | 0 | 20,981,010 | /s/ Andrew D. Dickinson | Andrew D. Dickinson, Executive Vice President and Chief Financial Officer | 0000882095 |
| Gilead Therapeutics A1 Unlimited Company | 29.9% | 20,981,010 | 0 | 20,981,010 | /s/ Aoife Marrinan | Aoife Marrinan, Director |