| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| Caligan Partners LP | 6.4% | 4,138,943 | 0 | 4,138,943 | /s/ David Johnson | David Johnson, Managing Partner | |
| David Johnson | 6.4% | 4,138,943 | 0 | 4,138,943 | /s/ David Johnson | David Johnson, Individually |