| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| American Century Investment Management, Inc. | 5.2% | 295,076 | 288,447 | 0 | American Century Investment Management, Inc. | /s/ John Pak / Senior Vice President | |
| American Century Companies, Inc. | 5.2% | 295,076 | 288,447 | 0 | American Century Companies, Inc. | /s/ John Pak / Senior Vice President | |
| Stowers Institute for Medical Research | 5.2% | 295,076 | 288,447 | 0 | Stowers Institute for Medical Research | /s/ Richard W. Brown / Chairman |